Very clever and satisfying. I found this one quite tricky, but steady work paid off. The way the titles replace is neatly done. It’s a bit of a pity the down “clue” answer doesn’t have a significance I can spot, but there’s so much else going on I can hardly complain! Thank you lath.
Just one parsing I can’t crack. I have the answer, as it’s all checked letters, and it fits the definition, and I know what letter is extra, but I just can’t figure out the wordplay. If anyone can nudge me to see it that will just close off the effort :-) It’s 22d, “gentleman rises to hug mostly top birds”.