I thought it was PLUMB early on and checked the various meanings in the dictionary but it was pushed out of the picture with BEETLE which gave a middle E instead of a middle U.
Incidentally, BEETLE is a short mallet-type thing for pounding, BEETLE is a game and BEETLE-headed means simple, stupid etc. So BEETLE seemed to stake a strong legitimate claim. And that was the problem and the obstacle that made the rest impossible to solve!