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22ac "Timid miaul with lion sadly turning into much tamer feline?" (13) Little to go on, but I'm new to this particular cryptic & it's taking time to get accustomed! Thanks in advance.

????????I???S

PUSILLANIMOUS
PUSS+ anagram of 'miaul' and 'lion'

Thank you very much. I was barking up completely the wrong tree & don't think I would ever had found the solution!